Esta región, que se extiende desde los bosques de tierras bajas de El Petén hasta los secos trópicos del valle del río Motagua, y desde el extremo del altiplano occidental hasta el mar Caribe, es la más variada de Guatemala.
Incluye: Salama y alrededores, Biotopo del Quetzal, Cobán, Lanquín, Chisec y alrededores, Chiquimula, Esquipulas, Quiriguá, Copán (Honduras), Lago de Izabal y Lívingston.
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